A. WARÉN. Murchisonellidae: who are they, where are they and what are they doing? (Gastropoda, lowermost Heterobranchia)
Abstract
The family Murchisonellidae is discussed to set it into its context among the Gastropoda and to add a dimension to Peñas & Rolán’s subsequent review of the species of the subfamily Murchisonellinae.
It is concluded that the name Ebalidae Warén, 1995 was validly introduced and worth at least subfamily status as a sister group to Murchisonellinae. The family name Anisocyclidae van Aartsen, 1995 is considered to have a type species closely related to that of Eulimella (Pyramidellidae: Eulimellinae).
Recent molecular work dates the separation of Murchisonella and Ebala back to the Carboniferous which may be an underestimate since it is likely that the earliest appearance of Heterobranchia still is unknown. In the discussions about other similar cases, Graphis is formally transferred from Aclididae (Caenogastropoda: Neotaenioglossa) and placed among the lower Heterobranchia in the family Graphididae, in accordance with recent molecular work. Tofanellidae is possibly an older name for Graphididae.
It is concluded that Murchisonellidae is probably the oldest, safely recognised, still living gastropod family. The recent species occur in fairly shallow water, 0-50 m depth and nothing is known about their nutritional ecology or what they use their modified radula for. The characteristic protoconch is inferred to be the result of planktotrophic development in most of the species.
A. PEÑAS & E. ROLÁN. Revision of the genera Murchisonella and Pseudoaclisina (Gastropoda, Heterobranchia, Murchisonellidae)
Abstract
A revision of the species of two genera of the family Murchisonellidae Casey, 1904, which have Recent representatives: Murchisonella Casey, 1904 and Pseudoaclisina Yoo, 1994, is presented. All the known species are figured, their morphologies described and comparisons made. In the first genus, Murchisonella, 22 species are recognised, from which 10 are new; in the other genus, Pseudoaclisina, there are 7 which all are new species for science.
New taxa
Murchisonella africana Peñas & Rolán, 2013
Type locality: Cape Verde Archipelago, Santiago Island, Tarrafal at 15 m
Murchisonella saotomensis Peñas & Rolán, 2013
Type locality: Minerio, 35-41 m, in sandy sediments, São Tomé Island
Murchisonella angolensis Peñas & Rolán, 2013
Type locality: Angola, Luanda, Corimba at 0-1 m
Murchisonella mediterranea Peñas & Rolán, 2013
Type locality: Turkey, Tasaçu, near Silifke
Murchisonella galapagensis Peñas & Rolán, 2013
Type locality: Islas Galápagos, San Salvador (James Island), Sullivan Bay, 36 m, (0º16’S, 90º34’W), Ecuador
Murchisonella curvistriata Peñas & Rolán, 2013
Type locality: Lagon du Grand Récif Mangalia (20°44’S 165°15.9’E), EXPÉDITION MONTROUZIER, Stn 1264, New Caledonia
Murchisonella latisulcata Peñas & Rolán, 2013
Type locality: Tamioka Bay, near Amakusa Marine Laboratory, Kumamoto Pref. (32º31.6’N, 130º03’E), Kyushu, 20-40 m, Japan
Murchisonella modesta Peñas & Rolán, 2013
Type locality: Pontod Islet, Panglao Island (9°33.6’N, 123°43.5’E), 0-3 m, PANGLAO 2004 Stn D5, Philippines
Murchisonella modestissima Peñas & Rolán, 2013
Type locality: Secteur de Touho, Lagon du Grand Récif Mangalia (20°44.5’S, 165°15.9’E), 8 m, LAGON Stn 1264, New Caledonia
Murchisonella dubia Peñas & Rolán, 2013
Type locality: Momo Beach (9°36.1’N, 123°45.2’E), 0-3 m, Panglao Is., PANGLAO 2004 Stn M7, Philippines
Pseudoaclisina indivisa Peñas & Rolán, 2013
Type locality: Secteur de Touho, Lagon du Grand Récif Mangalia (20°44.5’S, 165°15.9’E), 8 m EXPÉDITION MONTROUZIER Stn 1264, New Caledonia
Pseudoaclisina planicula Peñas & Rolán, 2013
Type locality: Secteur de Touho, Baie de Touho (20°46.7’S, 165°13.7’E), 3-6 m, EXPÉDITION MONTROUZIER Stn 1250, New Caledonia
Pseudoaclisina linealis Peñas & Rolán, 2013
Type locality: Secteur de Touho, Lagon du Grand Récif de Mangalia (20°44.5’S, 165°15.9’E), 8 m, EXPÉDITION MONTROUZIER Stn 1264, New Caledonia
Pseudoaclisina micrometrica Peñas & Rolán, 2013
Type locality: Secteur de Touho, Haut-Fond de Tié (20°52.7’N, 165°19.5’E), 5-25 m, EXPÉDITION MONTROUZIER Stn 1271, New Caledonia
Pseudoaclisina multistriata Peñas & Rolán, 2013
Type locality: Secteur de Koumac, Grand Récif de Koumac (20°41.4’S, 164°14.8’E), 20-30 m, EXPÉDITION MONTROUZIER Stn 1318, New Caledonia
Pseudoaclisina rarisulcata Peñas & Rolán, 2013
Type locality: Horseshoe Cliffs, 1 km WNW Onna Village, Okinawa (26º29.6’N, 127º50.5’E), Japan
Pseudoaclisina conica Peñas & Rolán, 2013
Type locality: Panglao, Sungcolan Bay (9°38.5’S, 123°49.7’E), 3-5 m, PANGLAO 2004 Stn B18, Philippines
